FLEUR publications

Submitted papers

  1. Caron MM, Vangansbeke P, Zellweger F, Verheyen K, Baeten L, Hedl R, Bernhardt-Romermann M, ...., De Frenne P (2024) Unequal community thermophilization in adult versus juvenile trees in European forests.
  2. De Lombaerde E, Vangansbeke P, Baeten L, Wei L, Landuyt D, Sanczuk P, Verheyen K, Bernhardt-Romermann M, ..., De Frenne P (2024) Thermophilization is triggering plant functional trait shifts in European temperate forests.
  3. Mazalla L, Fischer FM, Bello F, Biurrun I, Borovyk D, Brooker R, Brunet J, Campos JA, Carbognani M, Collins S, Danihelka J, De Frenne P, Pillar V, ..., Zobel M, Diekmann M (2024) Baseline turnover: accounting for stochasticity in studies of vegetation change.
  4. Sanczuk P, Verheyen K, Lenoir J, Zellweger F, Lembrechts J, Rodriguez-Sanchez F, Baeten L, Bernhardt-Romermann M, De Pauw K, ..., De Frenne P (2024) Unexpected east-west range shifts in European forest plants.
  5. Vanneste T, Graae B, Kyrkjeeide MO, Lindmo S, Michelsen O, Naranjo DJ, Ray CA, Vandersteene M, Verheyen K, De Frenne P (2024) Combining taxonomic, functional and phylogenetic diversity metrics to explain spatiotemporal variation in mountaintop flora.

Published or in press

  1. Haesen S, Lenoir J, Gril E, De Frenne P, Lembrechts J, Kopecky M, Macek M, Man M, Wild J, Van Meerbeek K (2023) Microclimate reveals the true thermal niche of forest plant species. Ecology Letters, 26, 2021-2166.

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  2. De Pauw K, Depauw L, Cousins SAO, De Lombaerde E, Diekmann M, Frey D, Kwietniowska K, Lenoir J, Meeussen C, Orczewska A, Plue J, Spicher F, Vanneste T, Zellweger F, Verheyen K, Vangansbeke P, De Frenne P (2024) The urban heat island accelerates litter decomposition through microclimatic warming in temperate urban forests. Urban Ecosystems, in press.

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  3. De Schuyter W, Depauw L, De Lombaerde E, De Smedt P, ..., Baeten L, De Frenne P, Bernhardt-Romermann M, Hedl R, Waller D, Verheyen K (2024) Declining potential nectar production of the herb layer in temperate forests under global change. Journal of Ecology, in press.

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  4. Huang S, Feigs JT, Holzhauer SIJ, Kramp K, Brunet J, Decocq G, De Frenne P, Diekmann M, Liira J, Spicher F, Vangansbeke P, Vanneste T, Verheyen K, Naaf T (2024) Limited effects of population age on the genetic structure of spatially isolated forest herb populations in temperate Europe. Ecology & Evolution, in press.
  5. Vanneste T, Depauw L, De Lombaerde E, Meeussen C, Govaert S, De Pauw K, ... De Frenne P (2024) Trade-offs of biodiversity and ecosystem services in European forest edges vs interiors. Nature Ecology & Evolution, in press.

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  6. Amstutz A, Firth LB, Spicer JI, De Frenne P, Gomez-Aparicio L, Graae BJ, Kus S, Lindmo S, Orczewska A, Rodriguez-Sanchez F, Vanneste T, Hanley M (2024) Taking sides? Aspect has limited influence on soil environment or litter decomposition in Pan-European study of roadside verges. Pedobiologia, 102, 150927.

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  7. De Pauw K, Depauw L, Cousins SAO, Decocq G, De Lombaerde E, Diekmann M, Frey D, Lenoir J, Meeussen C, Orczewska A, Plue J, Spicher F, Zellweger F, Vangansbeke P, Verheyen K, De Frenne P (2024) Nutrient-demanding and thermophilous plants dominate urban forest edge vegetation across temperate Europe. Journal of Vegetation Science, 35, e13236.

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  8. Govaert S, Meeussen C, Vanneste T, Bollmann K, Brunet J, Calders K, Cousins SAO, De Pauw K, Diekmann M, Graae BJ, Hedwall PO, Iacopetti G, Lenoir J, Lindmo S, Orczewska A, Ponette Q, Plue J, Sanczuk P, Selvi F, Spicher F, Verheyen K, Vangansbeke P, De Frenne P (2024) Trait - micro-environment relationships of forest herb communities across Europe. Global Ecology & Biogeography, 33, 286-302.

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  9. Moreira X, Abdala-Roberts L, Lago-Nunez B, Cao A, De Pauw K, De Ro A, Gasperini C, Hedwall PO, Iacopetti G, Lenoir J, Meeussen C, Plue J, Sanczuk P, Selvi F, Spicher F, Vanden Broeck A, Vangansbeke P, De Frenne P (2024) Effects of experimental warming at the microhabitat scale on oak leaf traits and insect herbivory across a contrasting environmental gradient. Oikos, 2024, e10353.

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  10. Padulles Cubino J, Lenoir J, Li D, Montano-Centellas F, Retana J, Baeten L, Bernhardt-Romermann M, Chudomelova M, Closset D, Decocq G, De Frenne P, Diekmann M, Dirnbock T, Durak T, Hedl R, Heinken T, Jaroszewicz B, Kopecky M, Macek M, Malis F, Naaf T, Orczewska A, Petrik P, Pielech R, Reczynska K, Schmidt W, Standovar T, Swierkosz K, Teleki B, Verheyen K, Vild O, Waller D, Wulf M, Chytry M (2024) Evaluating plant lineage losses and gains in temperate forest understories: a phylogenetic perspective on climate change and nitrogen deposition. New Phytologist, 241, 2287-2299.

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  11. De Pauw K, Depauw L, Caluwaerts C, Cousins SAO, De Lombaerde E, Diekmann M, Frey D, Lenoir J, Meeussen C, Orczewska A, Plue J, Spicher F, Zellweger F, Vangansbeke P, Verheyen K, De Frenne P (2023) Urban forest microclimates across temperate Europe are shaped by deep edge effects and forest structure. Agricultural and Forest Meteorology, 341, 109632.

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  12. Diaz-Calafat J, Uria-Diez J, Brunet J, De Frenne P, Vangansbeke P, Felton A, Ockinger E, Cousins SAO, Bauhus J, Ponette Q, Hedwall PO (2023) From broadleaves to conifers: The effect of tree composition and density on understory microclimate across latitudes. Agricultural & Forest Meteorology, 341, 109684.

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  13. Sanczuk P, De Pauw K, De Lombaerde E, Luoto M, Meeussen C, Govaert S, Vanneste T, Depauw L, Brunet J, Cousins SAO, Gasperini C, Hedwall PO, Iacopetti G, Lenoir J, Plue J, Selvi F, Spicher F, Uria-Diez J, Verheyen K, Vangansbeke P, De Frenne P (2023) Microclimate and forest density drive plant population dynamics under climate change. Nature Climate Change, 13, 840-847.

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  14. Wei L, Liira J, Ehrmann S, Lenoir J, Decocq G, Brunet J, Wulf M, Diekmann M, Naaf T, Scherer-Lorenzen M, Hansen K, De Smedt P, Valdes A, Verheyen K, De Frenne P (2023) Impact of patch age and size on forest soil characteristics in European agricultural landscapes. Science of the Total Environment, 898, 165543.

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  15. Wei L, Sanczuk P, De Pauw K, Caron MM, Selvi F, Hedwall PO, Brunet J, Cousins SAO, Plue J, Spicher F, Gasperini C, Iacopetti G, Orczewska A, Uria-Diez J, Lenoir J, Vangansbeke P, De Frenne P (2023) Using warming tolerances to predict understory plant responses to climate change. Global Change Biology, 30, e17064.

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  16. Auffret A, Vangansbeke P, De Frenne P, Auestad I, Basto S, Grandin U, Jacquemyn H, Jakobsson A, Kalamees R, Koch M, Marrs R, Marteinsdottir B, Wagner M, Bekker R, Bruun HH, Decocq G, Hermy M, Jankowska-Blaszczuk M, Milberg P, Maren I, Pakeman R, Phoenix G, Thompson K, Vandvik V, Plue J (2023) More warm-adapted species in soil seed banks than in herb-layer plant communities across Europe. Journal of Ecology, 111, 1009-1020.

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  17. De Lombaerde E, Vangansbeke P, Lenoir J, Van Meerbeek K, Lembrechts J, Rodriguez-Sanchez F, Luoto M, Scheffers B, Haesen S, Aalto J, Christiansen DM, De Pauw K, Depauw L, Govaert S, Greiser C, Hampe A, Hylander K, Klinges D, Koelemeijer I, Meeussen C, Ogee J, Sanczuk P, Vanneste T, Zellweger F, Baeten L, De Frenne P (2022) Maintaining forest cover to enhance temperature buffering under future climate change. Science of the Total Environment, 810, 151338.

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  18. Depauw L, Hu R, Hdungana K, Govaert S, Meeussen C, Vangansbeke P, Strimbeck, RG, Graae BJ, De Frenne P (2022) Functional trait variation of Anemone nemorosa along macro- and microclimatic gradients close to the northern range edge. Nordic Journal of Botany, 4, e03471.

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  19. De Pauw K, Sanczuk P, Meeussen C, Depauw L, De Lombaerde E, Govaert S, Vanneste T, Brunet J, Cousins SAO, Gasperini C, Hedwall PO, Jacopetti G, Lenoir J, Plue J, Selvi F, Spicher F, Uria-Diez J, Verheyen K, Vangansbeke P, De Frenne P (2022) Forest understorey communities respond strongly to light in interaction with forest structure, but not to microclimate warming. New Phytologist, 233, 219-235.

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  20. Fernandez PF, Sanczuk P, Vanneste T, Brunet J, Ehrlen J, Hedwall PO, Hylander K, Van Den Berge S, Verheyen K, De Frenne P (2022) Different effects of warming treatments in forests versus hedgerows on the understorey plant Geum urbanum. Plant Biology, 24, 734-744.

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  21. Gasperini C, Vanneste T, Bollmann K, Brunet J, Cousins SAO, Decocq G, De Pauw K, Diekmann M, Govaert S, Graae BJ, Hedwall PO, Iacopetti G, Lenoir J, Lindmo S, Meeussen C, Orczewska A, Ponette Q, Plue J, Sanczuk S, Spicher F, Vangansbeke P, Zellweger F, Selvi F, De Frenne P (2022) Soil seed bank responses to edge effects in temperate European forests. Global Ecology and Biogeography, 31, 1877-1893.

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  22. Hagenberg L, Vanneste T, Opedal O, Petlund H, Bjorkman M, Bjork R, Holien H, Limpens J, Molau U, Graae BJ, De Frenne P (2022) Vegetation change on mountaintops in northern Sweden: stable vascular-plant but reordering of lichen and bryophyte communities. Ecological Research, 37, 722--737.

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  23. Heinken T, Diekmann M, Liira J, Orczewska A, Schmidt M, Brunet J, Chytry M, Chabrerie O, Decocq G, De Frenne P, Drevojan P, Dzwonko Z, Ewald J, Feilberg J, Graae BJ, Grytnes JA, Hermy M, Kriebitzsch WU, Laivins M, Lenoir J, Lindmo S, Marage D, Marozas V, Niemeyer T, Paal J, Pysek P, Roosaluste E, Sadlo J, Schaminee J, Tyler T, Verheyen K, Wulf M, Vanneste T (2022) The European Forest Plant Species List (EuForPlant): Concept and Application. Journal of Vegetation Science, 33, e13132.

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  24. Lenoir J, Gril E, Durrieu S, Horen H, Laslier M, Lembrechts J, Zellweger F, Alleaume S, Brasseur B, Buridant J, Dayal K, De Frenne P, Gallet-Moron E, Marrec R, Van Meerbeek K, Meeussen C, Rocchini D, Decocq G (2022) Unveil the unseen: Using LiDAR to capture time-lag dynamics in the herbaceous layer of European temperate forests. Journal of Ecology, 110, 282--300.

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  25. Meeussen C, De Pauw K, Sanczuk P, Brunet J, Cousins SAO, Gasperini C, Hedwall PO, Iacopetti G, Lenoir J, Plue J, Selvi F, Spicher F, Diez JU, Verheyen K, Vangansbeke P, De Frenne P (2022) Initial oak regeneration responses to experimental warming along microclimatic and macroclimatic gradients. Plant Biology, 24, 745-757.

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  26. Naaf T, Feigs J, Huang S, Brunet J, Cousins SAO, Decocq G, De Frenne P, Diekmann M, Govaert S, Hedwall PO, Lenoir J, Liira J, Meeussen C, Plue J, Vangansbeke P, Vanneste T, Verheyen K, Holzhauer S, Kramp K (2022) Context matters: the landscape matrix determines the population genetic structure of temperate forest herbs across Europe. Landscape Ecology, 37, 1365-1384.

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  27. Segar JSR, Pereira H, Baeten L, Bernhardt-Romermann M, De Frenne P, Fernandez N, Gilliam F, Lenoir J, ... Wulf M, Staude I (2022) Divergent roles of herbivory in eutrophying forests. Nature Communications, 13, 7837.

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  28. Staude IR, Pereira HM, Daskalova G, ...Baeten L (2022) Directional turnover toward larger-ranged plants over time and across habitats. Ecology Letters, 25, 466-482.

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  29. Meeussen C, Govaert S, Vanneste T, Bollmann K, Brunet J, Calders K, Cousins SAO, De Pauw K, Diekmann M, Gasperini C, Hedwall PO, Hylander K, Iacopetti G, Lenoir J, Lindmo S, Orczewska A, Ponette Q, Plue J, Sanczuk P, Selvi F, Spicher F, Verbeeck H, Zellweger F, Verheyen K, Vangansbeke P, De Frenne P (2021) Microclimatic edge-to-interior gradients of European deciduous forests.Agricultural and Forest Meteorology, 311, 108699.

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  30. Freigs JT, Holzhauer SIJ, Huang S, Brunet J, Diekmann M, Hedwall PO, Kramp K, Naaf T (2022) Pollinator movement activity influences genetic diversity and differentiation of spatially isolated populations of clonal forest herbs. Frontiers in Ecology and Evolution, 10, 908258.

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  31. Caron MM, Zellweger F, Verheyen K, Baeten L, Hedl R, Bernhardt-Romermann M, Berki I, Brunet J, Decocq G, Diaz S, Dirnbock T, Durak T, Heinken T, Jaroszewicz B, Kopecky M, Lenoir J, Macek M, Marek M, Malis F, Nagel T, Perring M, Petrik P, Reczynska K, Pielech R, Schmidt W, Swierkosz K, Balazs T, Wulf M, De Frenne P (2022) Thermal differences between juveniles and adults increased over time in European forest trees. Journal of Ecology, 109, 3944-3957.

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  32. De Frenne P, Lenoir J, Luoto M, Scheffers B, Zellweger F, Aalto J, Ashcroft M, Christiansen D, Decocq G, De Pauw K, Govaert S, Greiser C, Gril E, Hampe A, Jucker T, Klinges D, Koelemeijer I, Lembrechts J, Marrec R, Meeussen C, Ogee J, Tyystjarvi V, Vangansbeke P, Hylander K (2021) Forest microclimates and climate change: importance, drivers and future research agenda. Global Change Biology, 27, 2279-2297.

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  33. Marsman F, Nystuen KO, Opedal OH, Foest JJ, Sorensen MV, De Frenne P, Graae BJ, Limpens J (2021) Determinants of pine tree establishment in alpine tundra. Journal of Vegetation Science, 32, e12948.

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  34. De Pauw K, Meeussen C, Govaert S, Sanczuk P, Vanneste T, Bernhardt-Romermann M, Bollmann K, Brunet J, Calders K, Cousins SAO, Diekmann M, Hedwall PO, Iacopetti G, Lenoir J, Lindmo S, Orczewska A, Ponette Q, Plue J, Selvi F, Spicher F, Verbeeck H, Vermeir P, Zellweger F, Verheyen K, Vangansbeke P, De Frenne P (2020) Taxonomic, phylogenetic and functional diversity of forest understorey plants along edge-to-interior gradients across Europe. Journal of Ecology, 109, 2629-2648.

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  35. Vanneste T, Van Den Berge S, Brunet J, Hedwall PO, Verheyen K, De Frenne P (2021) Temperature effects on forest understorey plants in hedgerows: a combined warming and transplant experiment. Annals of Botany, 128, 315-327.

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  36. Vangansbeke P, Malis F, Hedl R, Chudomelova M, Vild O, Wulf M, Jahn U, Welk E, Rodriguez-Sanchez F, De Frenne P (2021) ClimPlant: realized climatic niches of vascular plants in European forest understoreys. Global Ecology and Biogeography, 30, 1183-1190.

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  37. Naaf T, Feigs J, Huang S, Brunet J, Cousins SAO, Decocq G, De Frenne P, Diekmann M, Govaert S, Hedwall PO, Helsen K, Lenoir J, Liira J, Meeussen C, Plue J, Poli P, Spicher F, Vangansbeke P, Vanneste T, Verheyen K, Holzhauer S, Kramp K (2021) Forest herbs' sensitivity to habitat fragmentation across species and landscapes. Landscape Ecology, 36, 2831-2848.

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  38. Helsen K, Diekmann M, Decocq G, De Pauw K, Govaert S, Graae BJ, Hagenblad J, Liira J, Orczewska A, Sanczuk P, Van Meerbeek K, De Frenne P (2021) Biological Flora of Central Europe: Impatiens glandulifera Royle. Perspectives in Plant Ecology, Evolution and Systematics, 50, 125609.

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  39. Brunet, J., Hedwall, P.-O., Lindgren, J. & Cousins, S.A.O (2021) Immigration credit of temperate forest herbs in fragmented landscapes – implications for restoration of habitat connectivity. Journal of Applied Ecology, 58, 2195-2206.

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  40. Ronan Marrec, Vincent Le Roux, Ludmilla Martin, Jonathan Lenoir, Jörg Brunet, Sara AO Cousins, Pallieter De Smedt, Marc Deconchat, Martin Diekmann, Steffen Ehrmann, Emilie Gallet‐Moron, Brice Giffard, Jaan Liira, Jessica Lindgren, Alicia Valdes, Kris Verheyen, Monika Wulf, Guillaume Decocq (2021) Multiscale drivers of carabid beetle (Coleoptera: Carabidae) assemblages in small European woodlands. Global Ecology and Biogeography, 30, 165-182.

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  41. Meeussen C, Govaert S, Vanneste T, Haesen S, Van Meerbeek K, Bollmann K, Brunet J, Calders K, Cousins SAO, Diekmann M, Graae BJ, Iacopetti G, Lenoir J, Orczewska A, Ponette Q, Plue J, Selvi F, Spicher F, Vedel-Sorensen M, Verbeeck H, Vermeir P, Verheyen K, Vangansbeke P, De Frenne P (2021) Drivers of carbon stocks in forest edges across Europe. Science of the Total Environment, 759, 143497.

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  42. Marsman F, Nystuen KO, Opedal OH, Foest JJ, Sorensen MV, De Frenne P, Graae BJ, Limpens J (2021) Determinants of pine tree seedling establishment in alpine tundra. Journal of Vegetation Science, 32, e12948.

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  43. Sanczuk P, Govaert S, Meeussen C, De Pauw K, Vanneste T, Depauw L, Moreira X, Schoelynck J, De Boevre M, De Saeger S, Bollmann K, Brunet J, Cousins SAO, Plue J, Diekmann M, Graae BJ, Hedwall PO, Jacopetti G, Lenoir J, Orczewska A, Ponette Q, Selvi F, Spicher F, Vermeir P, Calders K, Verbeeck H, Verheyen K, Vangansbeke P, De Frenne P (2021) Small scale environmental variation modulates plant defence syndromes of understorey plants in deciduous forests of Europe. Global Ecology and Biogeography, 30, 205-219.

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  44. Helsen K, Acharya K, De Kort H, Brunet J, Chabrerie O, Cousins SAO, De Frenne P, Hermy M, Verheyen K, Graae BJ, Pelabon C (2020) Earlier onset of flowering and increased reproductive allocation of an annual invasive plant in the north of its novel range. Annals of Botany, 126, 1005-1016.

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  45. Vanneste T, Van Den Berge S, Riske E, Brunet J, Decocq G, Diekmann M, Graae BJ, Hedwall PO, Lenoir J, Liira J, Lindmo S, Litza K, Naaf T, Orczewska A, Wulf M, Verheyen K, De Frenne P (2020) Hedging against biodiversity loss: forest herbs' performance in hedgerows across temperate Europe. Journal of Vegetation Science, 108, 1750-1774.

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  46. Plue J, Cousins SAO, De Pauw K, Diekmann M, Hagenblad J, Helsen K, Hermy M, Liira J, Orczewska A, Vanneste T, Wulf M, De Frenne P (2020) Biological Flora of the British Isles: Poa nemoralis L. Journal of Ecology, 108, 1750-1774.

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  47. Zellweger F, De Frenne P, Lenoir J, Vangansbeke P, Verheyen K, Bernhardt-Romermann M, Baeten L, Hedl R, Berki I, ... Wulf M, Coomes DA (2020) Forest microclimate dynamics drive plant responses to warming. Science, 368, 772-775.

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  48. Staude IR, Waller DM, Bernhardt-Romermann M, Bjorkman AD, Brunet J, .... Baeten L (2020) Replacements of small- by large-ranged species scale up to diversity loss in Europe's temperate forest biome. Nature Ecology & Evolution, 4, 802-808.

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  49. Vanneste T, Govaert S, De Kesel W, Van Den Berge S, Vangansbeke P, Meeussen C, Brunet J, Cousins SAO, Decocq G, Diekmann M, Graae BJ, Hedwall PO, Heinken T, Helsen K, Kapas R, Lenoir J, Liira J, Lindmo S, Litza K, Naaf T, Orczewska A, Plue J, Wulf M, Verheyen K, De Frenne P (2020) Plant diversity in hedgerows and road verges across Europe. Journal of Applied Ecology, 57, 1244-1257.

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  50. Meeussen C, Govaert S, Vanneste T, Calders K, Bollmann K, Brunet J, Cousins SAO, Diekmann M, Graae BJ, Hedwall PO, Krishna Moorthy SM, Iacopetti G, Lenoir J, Lindmo S, Orczewska A, Ponette Q, Plue J, Selvi F, Spicher F, Tolosano M, Verbeeck H, Verheyen K, Vangansbeke P, De Frenne P (2020) Structural variation of forest edges across Europe. Forest Ecology and Management, 462, 117929.

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  51. Govaert S, Meeussen C, Vanneste T, Bollmann K, Brunet J, Cousins SAO, Diekmann M, Graae BJ, Hedwall PO, Iacopetti G, Lenoir J, Lindmo S, Orczewska A, Perring M, Ponette Q, Plue J, Selvi F, Spicher F, Tolosano M, Vermeir P, Zellweger F, Verheyen K, Vangansbeke P, De Frenne P (2020) Edge influence on understorey plant communities depends on forest management. Journal of Vegetation Science, 31, 281-292.

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  52. Valdes A, Lenoir J, De Frenne P, Andrieu E, Brunet J, Chabrerie O, Cousins SAO, Deconchat M, De Smedt P, Diekmann M, Ehrmann S, Gallet-Moron E, Gartner S, Giffard B, Hansen K, Hermy M, Kolb A, Le Roux V, Liira J, Lindgren J, Martin L, Naaf T, Paal T, Proesmans W, Scherer-Lorenzen M, Wulf M, Verheyen K, Decocq G (2020) High ecosystem service delivery potential of small woodlands in agricultural landscapes. Journal of Applied Ecology, 57, 4-16.

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  53. Vanneste T, Govaert S, Spicher F, Brunet J, Cousins SAO, Decocq G, Diekmann M, Graae BJ, Hedwall PO, Kapas R, Lenoir J, Liira J, Lindmo S, Litza K, Naaf T, Orczewska A, Plue J, Wulf M, Verheyen K, De Frenne P (2020) Contrasting microclimates among hedgerows and woodlands across temperate Europe. Agricultural and Forest Meteorology, 281, 107818.

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  54. Zellweger F, Coomes D, Lenoir J, Depauw L, Maes S, Wulf M, Kirby K, Brunet J, Kopecky M, Malis F, Schmidt W, Heinrichs S, den Ouden J, Jaroszewicz B, Buyse G, Spicher F, Verheyen K, De Frenne P (2019) Seasonal drivers of understorey temperature buffering in temperate deciduous forests across Europe. Global Ecology and Biogeography, 28, 1774-1786.

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  55. De Frenne P, Zellweger F, Rodriguez-Sanchez F, Scheffers B, Hylander K, Luoto M, Vellend M, Verheyen K, Lenoir J (2019) Global buffering of temperatures under forest canopies. Nature Ecology and Evolution, 3, 744-749.

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  56. Ma S, De Frenne P, Boon N, Brunet J, Cousins SAO, Decocq G, Kolb A, Lemke I, Liira J, Naaf T, Orczewska A, Plue J, Wulf M, Verheyen K (2019) Plant species identity and soil characteristics determine rhizosphere soil bacteria community composition in European temperate forests. FEMS Microbiology Ecology, 95, fiz063.

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  57. Proesmans W, Bonte D, Smagghe G, Meeus I, Decocq G, Spicher F, Kolb A, Lemke I, Diekmann M, Bruun HH, Wulf M, Van Den Berge S, Verheyen K (2019) Small forest patches as pollinator habitat: oases in an agricultural desert? Landscape Ecology, 34, 487-501.

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  58. Zellweger F, De Frenne P, Lenoir J, Rocchini D, Coomes DA (2019) Advances in microclimate ecology arising from remote sensing. Trends in Ecology and Evolution, 34, 327-341.

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  59. Ma S, De Frenne P, Vanhellemont M, Wasof S, Boeckx P, Brunet J, Cousins SAO, Decocq G, Kolb A, Lemke I, Liira J, Naaf T, Orczewska A, Plue J, Wulf M, Verheyen K (2019) Local soil characteristics determine the microbial communities under forest understorey plants along a latitudinal gradient. Basic and Applied Ecology, 36, 34-44.

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  60. De Smedt P, Baeten L, Proesmans W, Van de Poel S, Van Keer J, Giffard B, Martin L, Vanhulle R, Brunet, J, Cousins SAO, Decocq G, Deconchat M, Diekmann M, Gallet-Moron E, Le Roux V, Liira J, Valdes A, Wulf M, Andrieu E, Hermy M, Bonte D, Verheyen K (2019) Strength of forest edge effects on litter-dwelling macroarthropods across Europe is influenced by forest age and edge properties. Diversity and distributions, 25, 963-974.

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  61. Helsen K, Hagenblad J, Acharya KP, Brunet J, Cousins SAO, Decocq G, De Frenne P, Kimberley A, Kolb A, Michaelis J, Plue J, Verheyen K, Speed JDM, Graae BJ (2019) No genetic erosion after five generations for Impatiens glandulifera populations across the invaded range in Europe. BMC Genetics, 20, 20.

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  62. Ma S, De Frenne P, Vanhellemont M, Wasof S, Brunet J, Cousins SAO, Decocq G, Kolb A, Lemke I, Liira J, Naaf T, Orczewska A, Plue J, Wulf M, Verheyen K (2019) Plant-soil feedbacks of forest understorey plants transplanted in nonlocal soils along a latitudinal gradient. Plant Biology, 21, 677-687.

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  63. Verheyen K, Bazany M, Checko E, Chudomelova M, Closset-Kopp D, Czortek P, Decocq G, De Frenne P, De Keersmaeker L, Enriquez-Garcia C, Fabsicova M, Grytnes JA, Hederova L, Hedl R, Heinken T, ..., Wulf M, Baeten L (2018) Observer and relocation errors matter in resurveys of historical vegetation plots. Journal of Vegetation Science, 29, 812-823.

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  64. Vanneste T, Valdes A, Verheyen K, Perring MP, Bernhardt-Romermann M, Andrieu E, Brunet J, Cousins SAO, Deconchat M, De Smedt P, Diekmann M, Ehrmann S, Heinken T, Hermy M, Kolb A, Lenoir J, Liira J, Naaf T, Paal T, Wulf M, Decocq G, De Frenne P. Forest plant functional trait variation across Europe. Basic and Applied Ecology, 34, 1-14.

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  65. Perring MP, Diekmann M, Midolo G, Schellenberger Costa D, Bernhardt-Romermann M, Otto J, Gilliam F, Hedwall PO, Nordin A, Dirnbock T, Simkin S, Malis F, Blondeel H, Brunet J, Chudomelova M, Durak T, De Frenne P, Hedl R, Kopecky M, Landuyt D, Li D, Manning P, Petrik P, Reczynska K, Schmidt W, Standovar T, Swierkosz K, Vild O, Waller D, Verheyen K (2018) Understanding context dependency in the response of forest understorey plant communities to nitrogen deposition. Environmental Pollution, 242, 1787-1799.

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  66. De Smedt P, Baeten L, Proesmans W, Berg MP, Brunet J, Cousins SAO, Decocq G, Deconchat M, Diekmann M, Gallet-Moron E, Giffard B, Liira J, Martin L, Ooms A, Valdes A, Wulf M, Hermy M, Bonte D, Verheyen K (2018) Linking macrodetritivore distribution to desiccation resistance in small forest fragments embedded in agricultural landscapes in Europe. Landscape Ecology, 33, 407-421.

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  67. De Frenne P, Blondeel H, Brunet J, Caron MM, Chabrerie O, Cougnon M, Cousins SAO, Decocq G, Diekmann M, Graae BJ, Hanley ME, Heinken T, Hermy M, Kolb A, Lenoir J, Liira J, Orczewska A, Shevtsova A, Vanneste T, Verheyen K (2018) Atmospheric nitrogen deposition on petals enhances seed quality of the forest herb Anemone nemorosa. Plant Biology, 20, 619-626.

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  68. Helsen K, Smith S, Brunet J, Cousins SAO, De Frenne P, Kolb A, Lenoir J, Ma S, Michaelis J, Plue J, Verheyen K, Speed J, Graae BJ (2018) Impact of an invasive alien plant on litter decomposition along a latitudinal gradient. Ecosphere, 9, e02097.

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  69. Perring MP, Bernhardt-Romermann M, Baeten L, Blondeel H, Depauw L, Landuyt D, Maes SL, De Lombaerde E, Caron MM, Vellend M, Brunet J, Chudomelova M, Decocq G, Diekmann M, Dirnbock T, Dorfler I, Durak T, De Frenne P, Gilliam F, Hedl R, Heinken T, Jaroszewicz B, Kirby KJ, Kopecky M, Lenoir J, Li D, Malis F, Naaf T, Newman M, Petrik P, Reczynska K, Schmidt W, Standovar T, Swierkosz K, Van Calster H, Vild O, Wagner ER, Wulf M, Verheyen K (2018) Global environmental change effects on plant community composition trajectories depend upon management legacies. Global Change Biology, 24, 1722-1740.

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  70. Ehrmann S, Ruyts SC, Scherer-Lorenzen M, Brunet J, Cousins SAO, Deconchat M, Decocq G, De Frenne P, De Smedt P, Diekmann M, Gallet-Moron E, Gartner S, Hansen K, Kolb A, Lenoir J, Lindgren J; Naaf T, Paal T, Panning M, Prinz M, Valdes A, Verheyen K, Liira J (2018) Habitat properties are key drivers of Borrelia burgdorferi s.l. prevalence in Ixodes ricinus populations of deciduous forest fragments. Parasites & Vectors, 11, 23.

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  71. De Smedt P, Baeten L, Berg MP, Gallet-Moron E, Brunet J, Cousins SAO, Decocq G, Diekmann M, Giffard B, De Frenne P, Hermy M, Bonte D, Verheyen K (2018) Desiccation resistance determines distribution of woodlice along forest edge-to-interior gradients. European Journal of Soil Biology, 85, 1-3.

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  72. Helsen K, Acharya K, Brunet J, Cousins SAO, Decocq G, Hermy M, Kolb A, Lemke I, Lenoir J, Plue J, Verheyen K, De Frenne P, Graae BJ (2017) Biotic and abiotic drivers of intraspecific trait variation within plant populations of three herbaceous plant species along a latitudinal gradient. BMC Ecology, 17, 38.

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  73. Ehrmann S, Liira J, Gartner S, Hansen K, Bauhus J, Brunet J, Cousins SAO, Deconchat M, Decocq G, De Frenne P, De Smedt P, Diekmann M, Gallet-Moron E, Kolb A, Lenoir J, Lindgren J, Naaf T, Paal T, Valdes A, Verheyen K, Wulf M, Scherer-Lorenzen M (2017) Environmental drivers of Ixodes ricinus abundance in forest fragments of rural European landscapes. BMC Ecology,17, 31.

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  74. Vanneste T, Michelsen O, Graae BJ, Olsen Kyrkjeeide M, Holien H, Hassel K, Lindmo S, De Frenne P (2017) Impact of climate change on alpine vegetation of mountain summits in Norway. Ecological Research, 32, 579-593.

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  75. Acharya KP, De Frenne P, Brunet J, Chabrerie O, Cousins SAO, Diekmann M, Hermy M, Kolb A, Lemke I, Plue J, Verheyen K & Graae BJ (2017) Latitudinal variation of life-history traits of an exotic and a native Impatiens species in Europe. Acta Oecologica, 81, 40-47.

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  76. De Frenne P, Brunet J, Cougnon M, Decocq G, Graae BJ, Hagenblad J, Hermy M, Kolb A, Lemke I, Ma S, Orczewska A, Plue J, Vranckx G, Wulf M & Verheyen K. Biological Flora of the British Isles: Milium effusum. Journal of Ecology, 105, 839-858.

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  77. Plue J, De Frenne P, Acharya KP, Brunet J, Chabrerie O, Decocq G, Diekmann M, Graae BJ, Heinken T, Hermy M, Kolb A, Lemke I, Liira J, Naaf T, Verheyen K, Wulf M, Cousins SAO (2017) Where Does the Community Start, and Where does it End? Including the Seed Bank to Reassess Forest Herb Layer Responses to the Environment. Journal of Vegetation Science, 28, 424-435.

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  78. Verheyen K, De Frenne P, Baeten L, Waller D, Hedl R, Perring M, Brunet J, Chudomelova M, De Lombaerde E, Depauw L, Dirnboeck T, Durak T, Eriksson O, Gilliam F, Heinken T, Heinrichs S, Hermy M, Jaroszewicz B, Jenkins M, Johnson S, Kirby K, Kopecky M, Landuyt D, Lenoir J, Li D, Macek M, Maes SL, Malis F, Mitchell F, Naaf T, Peterken GP, Petrik P, Reczynska K, Rogers DA, Hoistad Schei F, Schmidt W, Standovar T, Swierkosz K, Ujhazy K, Van Calster H, Vellend M, Vild O, Woods K, Wulf M, Bernhardt-Romermann M (2017) Combining community resurvey data to advance global change research. BioScience, 67, 73-83.

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  79. Reinecke J, Wulf M, Baeten L, Brunet J, Decocq G, De Frenne P, Diekmann M, Graae BJ, Heinken T, Hermy M, Jamoneau A, Lenoir J, Plue J, Orczewska A, Van Calster H, Verheyen K, Naaf T (2016) Acido- and neutrophilic temperate forest plants display distinct shifts in ecological pH niche across north-western Europe. Ecography, 39, 1164-1175.

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  80. Decocq G, Andrieu E, Brunet J, Chabrerie O, De Frenne P, De Smedt P, Deconchat M, Diekmann M, Ehrmann S, Giffard B, Gorriz Mifsud E, Hansen K, Hermy M, Kolb A, Lenoir J, Liira J, Moldan F, Prokofieva I, Rosenqvist L, Varela E, Valdes A, Verheyen K & Wulf M (2016) Ecosystem services from small forest patches in agricultural landscapes. Current Forestry Reports, 2, 30-44.

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  81. Lemke IH, Kolb A, Graae BJ, De Frenne P, Acharya KP, Blandino C, Brunet J, Chabrerie O, Cousins SAO, Decocq G, Heinken T, Hermy M, Liira J, Schmucki R, Shevtsova A, Verheyen K & Diekmann M (2015) Patterns of phenotypic trait variation in two temperate forest herbs along a broad climatic gradient. Plant Ecology, 216, 1523-1536.

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  82. Bernhardt-Römermann M, Baeten L, Craven D, De Frenne P, Hedl R, Lenoir J, Bert D, Brunet J, Chudomelova M, Decocq G, Dierschke H, Dirnbock T, Dorfler I, Heinken T, Hermy M, Hommel P, Jaroszewicz B, Keczynski A, Kelly DL, Kirby KJ, Kopecky M, Macek M, Malis F, Mirtl M, Mitchell FJG, Naaf T, Newman M, Peterken G, Petrik P, Schmidt W, Standovar T, Toth Z, Van Calster H, Verstraeten G, Vladovic J, Vild O, Wulf M & Verheyen K (2015) Drivers of temporal changes in temperate forest plant diversity vary across spatial scales. Global Change Biology, 21, 3726-3737.

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  83. Hagenblad J, Hulskotter J, Acharya KP, Brunet J, Chabrerie O, Cousins SAO, Dar PA, Diekmann M, De Frenne P, Jamoneau A, Kolb A, Lemke I, Plue J, Reshi ZA & Graae BJ (2015) Low genetic diversity despite multiple introductions of the invasive plant species Impatiens glandulifera in Europe. BMC Genetics, 16, 103.

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  84. Valdes A, Lenoir J, Gallet-Moron E, Andrieu E, Brunet J, Chabrerie O, Closset-Kopp D, Cousins SAO, Deconchat M, De Frenne P, De Smedt P, Diekmann M, Hansen K, Hermy M, Kolb A, Liira J, Lindgren J, Naaf T, Paal T, Prokofieva I, Scherer-Lorenzen M, Wulf M, Verheyen K & Decocq G (2015) The contribution of patch-scale conditions is greater than that of macroclimate in explaining local plant diversity in fragmented forests across Europe. Global Ecology and Biogeography, 24, 1094-1105.

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  85. Carón MM, De Frenne P, Chabrerie O, Cousins SAO, De Backer L, Decocq G, Diekmann M, Heinken T, Kolb A, Naaf T, Plue J, Selvi F, Strimbeck GR, Wulf M & Verheyen K (2015) Impacts of warming and changes in precipitation frequency on the regeneration of two Acer species. Flora, 214, 24-33.

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  86. Carón MM, De Frenne P, Brunet J, Chabrerie O, Cousins SAO, Decocq G, Diekmann M, Graae BJ, Heinken T, Kolb A, Lenoir J, Naaf T, Plue J, Selvi F, Wulf M & Verheyen K (2015) Divergent regeneration responses of two closely related tree species to direct abiotic and indirect biotic effects of climate change. Forest Ecology and Management, 342, 21-29.

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  87. Carón MM, De Frenne P, Brunet J, Chabrerie O, Cousins SAO, De Backer L, Decocq G, Diekmann M, Heinken T, Kolb A, Naaf T, Plue J, Selvi F, Strimbeck R, Wulf M, & Verheyen K (2015) Interacting effects of warming and drought on regeneration and early growth of Acer pseudoplatanus and A. platanoides. Plant Biology, 17, 52-62.

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  88. Carón MM, De Frenne P, Brunet J, Chabrerie O, Cousins SAO, De Backer L, Diekmann M, Graae BJ, Heinken T, Kolb A, Naaf T, Plue J, Selvi F, Strimbeck GR, Wulf M & Verheyen K (2014) Latitudinal variation in seeds characteristics of Acer platanoides and A. pseudoplatanus. Plant Ecology, 215, 911-925.

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  89. Maes S, De Frenne P, Brunet J, de la Pena E, Chabrerie O, Cousins SAO, Decocq G, Diekmann M, Gruwez R, Hermy M, Kolb A & Verheyen K (2014) Effects of enhanced nitrogen inputs and climate warming on a forest understorey plant assessed by transplant experiments along a latitudinal gradient. Plant Ecology, 215, 899-910.

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  90. De Frenne P, Rodriguez-Sanchez F, Bernhardt-Römermann M, Brown CD, Eriksson O, Hermy M, Mitchell FJG, Petrik P, Van Calster H, Vellend M & Verheyen K (2014) Reply to Harwood et al.: Thermophilization estimation is robust to the scale of species distribution data. Proceedings of the National Academy of Sciences, USA (PNAS), 111, E1166.

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  91. De Frenne P, Coomes DA, De Schrijver A, Staelens J, Alexander JM, Bernhardt-Römermann M, Brunet J, Chabrerie O, Chiarucci A, den Ouden J, Eckstein RL, Graae BJ, Gruwez R, Hedl R, Hermy M, Kolb A, Marell A, Mullender SM, Olsen SL, Orczewska A, Peterken G, Petrik P, Plue J, Simonson WD, Tomescu CV, Vangansbeke P, Verstraeten G, Vesterdal L, Wulf M & Verheyen K (2014) Plant movements and climate warming: intraspecific variation in growth responses to non-local soils. New Phytologist, 202, 431-441.

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  92. Baeten L, Warton DI, Van Calster H, De Frenne P, Verstraeten G, Bonte D, Bernhardt-Römmerman M, Cornelis J, Decocq G, Eriksson O, Hedl R, Heinken T, Hermy M, Hommel P, Kirby K, Naaf T, Petrik P, Walther GR, Wulf M & Verheyen K (2014) A model-based approach to studying changes in compositional heterogeneity. Methods in Ecology and Evolution, 5, 156-164.

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  93. De Frenne P, Rodriguez-Sanchez F, Coomes DA, Baeten L, Verstraeten G, Vellend M, Bernhardt-Romermann M, Brown CD, Brunet J, Cornelis J, Decocq G, Dierschke H, Eriksson O, Gilliam FS, Hedl R, Heinken T, Hermy M, Hommel P, Jenkins MA, Kelly DL, Kirby KJ, Mitchell FJG, Naaf T, Newman M, Peterken G, Petrik P, Schulz J, Sonnier G, Van Calster H, Waller DM, Walther GR, White PS, Woods K, Wulf M, Graae BJ & Verheyen K (2013) Microclimate moderates plant responses to macroclimate warming. Proceedings of the National Academy of Sciences, USA (PNAS), 110, 18561-18565.

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  94. Wasof, S., Lenoir, J., Gallet-Moron, E., Jamoneau, A., Brunet, J., Cousins, S.A.O., De Frenne, P., Diekmann, M., Hermy, M., Kolb, A., Liira, J., Verheyen, K., Wulf, M. & Decocq, G. (2013) Ecological niche shifts of understory plants along a latitudinal gradient of temperate deciduous forests. Global Ecology and Biogeography, 22, 1130-1140.

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  95. Plue, J., De Frenne, P., Acharya, K.P., Brunet, J., Chabrerie, O., Decocq, G., Diekmann, M., Graae, B.J., Heinken, T., Hermy, M., Kolb, A., Lemke, I., Liira, J., Naaf, T., Shevtsova, A., Verheyen, K., Wulf, M. & Cousins, S.A.O. (2013) Climatic control of forest herb seed banks along a latitudinal gradient. Global Ecology and Biogeography, 22, 1106-1117.

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  96. De Frenne, P., Graae, B.J., Rodriguez-Sanchez, F., Kolb, A., Chabrerie, O., Decocq, G., De Kort, H., De Schrijver, A., Diekmann, M., Eriksson, O., Gruwez, R., Hermy, M., Lenoir, J., Plue, J., Coomes, D.A. & Verheyen K. (2013) Latitudinal gradients as natural laboratories to infer species' responses to temperature. Journal of Ecology, 101, 784-795.

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  97. Brunet, J., De Frenne, P., Holmstrom, E. & Mayr, L.M. (2012) Life-history traits explain rapid colonization of young post-agricultural forests by understory herbs. Forest Ecology and Management, 278, 55-62.

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  98. De Frenne, P., Graae, B.J., Brunet, J., Shevtsova, A., De Schrijver, A., Chabrerie, O., Cousins, S.A.O., Decocq, G., Diekmann, M., Hermy, M., Heinken, T., Kolb, A., Nilsson, C., Stanton, S. & Verheyen, K. (2012) The response of forest plant regeneration to temperature variation along a latitudinal gradient. Annals of Botany, 109, 1037-1046.

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  99. Verheyen, K., Baeten, L., De Frenne, P., Bernhardt-Römermann, M., Brunet, J., Cornelis, J., Decocq, G., Dierschke, H., Eriksson, O., Hedl, R., Heinken, T., Hermy, M., Hommel, P., Kirby, K., Naaf, T., Peterken, G., Petrik, P., Pfadenhauer, J., Van Calster, H., Walther, G.-R., Wulf, M. & Verstraeten, G. (2012) Driving factors behind the eutrophication signal in understorey plant communities of deciduous temperate forests. Journal of Ecology, 100, 352-365.

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  100. Graae, B.J., De Frenne, P., Kolb, A., Brunet, J., Chabrerie, O., Verheyen, K., Pepin, N., Heinken, T., Zobel, M., Shevtsova, A., Nijs, I. & Milbau, A. (2012) On the use of weather data in ecological studies along altitudinal and latitudinal gradients. Oikos, 121, 3-19.

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  101. De Frenne, P., Brunet, J., Shevtsova, A., Kolb, A., Graae, B.J., Chabrerie, O., Cousins, S.A.O., Decocq, G., De Schrijver, A., Diekmann, M., Gruwez, R., Heinken, T., Hermy, M., Nilsson, C., Stanton, S., Tack, W., Willaert, J., & Verheyen, K. (2011) Temperature effects on forest herbs assessed by warming and transplant experiments along a latitudinal gradient. Global Change Biology, 17, 3240-3253.

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  102. De Frenne, P., Kolb, A., Graae, B.J., Decocq, G., Baltora, S., De Schrijver, A., Brunet, J., Chabrerie, O., Cousins, S.A.O., Dhondt, R., Diekmann, M., Gruwez, R., Heinken, T., Hermy, M., Liira, J., Saguez, R., Shevtsova, A., Baskin, C.C. & Verheyen, K. (2011) A latitudinal gradient in seed nutrients of the forest herb Anemone nemorosa. Plant Biology, 13, 493-501.

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  103. De Frenne, P., Graae, B.J., Kolb, A., Shevtsova, A., Baeten, L., Brunet, J., Chabrerie, O., Cousins, S.A., Decocq, G., Dhondt, R., Diekmann, M., Gruwez, R., Heinken, T., Hermy, M., Oster, M., Saguez, R., Stanton, S., Tack, W., Vanhellemont, M. & Verheyen, K. (2011) An intraspecific application of the leaf-height-seed ecology strategy scheme to forest herbs along a latitudinal gradient. Ecography, 34, 132-140.

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  104. De Frenne, P., Baeten, L., Graae, B.J., Brunet, J., Wulf, M., Orczewska, A., Kolb, A., Jansen, I., Jamoneau, A., Jacquemyn, H., Hermy, M., Diekmann, M., De Schrijver, A., De Sanctis, M., Decocq, G., Cousins, S.A.O. & Verheyen, K. (2011) Interregional variation in the floristic recovery of post-agricultural forests. Journal of Ecology, 99, 600-609.

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  105. De Frenne, P., Graae, B., Kolb, A., Brunet, J., Chabrerie, O., Cousins, S., Decocq, G., Diekmann, M., Dhondt, R., Eriksson, O., Heinken, T., Hermy, M., Jogar, U., Saguez, R., Shevtsova, A., Stanton, S., Zindel, R., Zobel, M. & Verheyen, K. (2010) Significant effects of temperature on the reproductive output of the forest herb Anemone nemorosa L. Forest Ecology and Management, 259, 809-817.

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  106. De Frenne, P., Kolb, A., Verheyen, K., Brunet, J., Chabrerie, O., Decocq, G., Diekmann, M., Eriksson, O., Heinken, T., Hermy, M., Jogar, U., Stanton, S., Quataert, P., Zindel, R., Zobel, M. & Graae, B.J. (2009) Unravelling the effects of temperature, latitude and local environment on the reproduction of forest herbs. Global Ecology and Biogeography, 18, 641-651.

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  107. Graae, B.J., Verheyen, K., Kolb, A., Van der Veken, S., Heinken, T., Chabrerie, O., Diekmann, M., Valtinat, K., Zindel, R., Karlsson, E., Strom, L., Decocq, G., Hermy, M. & Baskin, C. (2009) Germination requirements and seed mass of slow and fast colonizing forest herbs along a latitudinal gradient. Ecoscience, 16, 248-257.

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